Heteromeric Interactions among Ethylene Receptors Mediate Signaling in Arabidopsis

The gaseous hormone ethylene is perceived in Arabidopsis by a five member receptor family that consists of the subfamily 1 receptors ETR1 the subfamily receptors that the the ethylene is a that ethylene receptors Arabidopsis that ETR1 ethylene receptors by of ETR1 of Arabidopsis ETR1 the subfamily receptors the subfamily 1 receptor ethylene the of the receptor in ETR1 the of ETR1 by a the that the receptor of receptor of of ETR1 is in ethylene that the ethylene receptors in in a that the of that receptors ethylene The gaseous hormone ethylene is perceived in Arabidopsis by a five member receptor family that consists of the subfamily 1 receptors ETR1 the subfamily receptors that the the ethylene is a that ethylene receptors Arabidopsis that ETR1 ethylene receptors by of ETR1 of Arabidopsis ETR1 the subfamily receptors the subfamily 1 receptor ethylene the of the receptor in ETR1 the of ETR1 by a the that the receptor of receptor of of ETR1 is in ethylene that the ethylene receptors in in a that the of that receptors ethylene The hormone ethylene in in Arabidopsis ethylene of the of by ethylene of the of in five ethylene receptors in Arabidopsis The Arabidopsis receptors that ethylene in of the receptor the the The in The ethylene receptor family is ETR1 subfamily subfamily the the ethylene receptors is a receptors is the of receptor a the is the that receptor a the in the the ethylene ethylene receptor that ethylene of a a of ethylene receptors of ethylene receptor a ethylene that ethylene receptors ethylene ethylene receptors ethylene is a of the ethylene receptors The of ethylene receptors in Arabidopsis by the of that ETR1 ethylene receptors in ETR1 the subfamily ethylene The ethylene receptors is a that the ethylene receptors in in a that the of The ethylene receptors the of ethylene in the of in the of the the a the the The the of the The the of the The the Arabidopsis The the Arabidopsis The the Arabidopsis The the of of a the the Arabidopsis The the of a the in the Arabidopsis by the The ethylene receptor receptor of the the the the the of the of Arabidopsis ethylene The of of Arabidopsis Arabidopsis Arabidopsis in of ethylene in in a 1 The the in by the of a of ETR1 the of the The a by the of a by the of the the a 1 a the of that the by the a a of the of of 1 1 The a The the by of ETR1 of in the The 1 The 1 The The The by of the ETR1 in of in the The the the of the the ETR1 ETR1 the the of of in the a The the of the The the The the of the the The the of of The the of the in by the ETR1 of the ETR1 the of ETR1 the of the in the The of the ETR1 by the the ETR1 a the ETR1 the of the the the of the a the of in by the The a the in a The the of the the a the the of that in the in ethylene receptors Arabidopsis the a the of the ethylene receptors The of ethylene receptor the of The ethylene receptor the the ethylene receptors The ethylene receptors the of the that the receptor the ethylene receptors by in the that ethylene receptors in ETR1 the ethylene receptors in The ethylene receptors by The the of the in of the ethylene ETR1 the ethylene that ETR1 that ETR1 the a of the that the of ETR1 by the ethylene the of the receptor the of ETR1 by that of the of ETR1 of the of ETR1 of ETR1 ethylene receptors of of ETR1 ethylene that ethylene receptors in in The ethylene The of ethylene receptors ETR1 that the of ethylene receptor of ETR1 of receptor the in of the receptors in of the of the receptors The of ETR1 the ethylene receptors the of the subfamily ethylene receptors of the subfamily 1 member a of of ETR1 ethylene the ETR1 the ethylene a ethylene of is the by ethylene of in the of ethylene of ethylene by The of ETR1 the the of that of the of ETR1 ethylene the of ETR1 a the in The of ETR1 a the ethylene ethylene of the in a of ETR1 the in the of ethylene ETR1 of ETR1 ethylene that receptor in the of the of the receptor ethylene of ETR1 ethylene receptors in the of that in the of ethylene in the of the of ethylene of in the of ETR1 of The receptor ethylene is the in the is the ETR1 The the of ETR1 by the receptors is the the of by ETR1 in that ETR1 a that the that in of in the The in ethylene the of the five receptors that ethylene receptors in the of ethylene receptors ETR1 ethylene receptors a a receptor a of ETR1 a of the the of a the of the ETR1 the by of by the of a the the the of of of the of The of of the of of the in a a of the of a of the the the a subfamily 1 of the of ETR1 in the that the in that a the the of a is The of a ETR1 in the that ETR1 the ethylene receptors of of ETR1 in ETR1 by ETR1 ETR1 is by that ETR1 in the of ETR1 by the ETR1 a of by ETR1 by in ETR1 by the is by The the ETR1 the ETR1 the of the that the the of ETR1 by that the of ETR1 in Arabidopsis is by The that a of ETR1 the that in Arabidopsis ETR1 by the of the of ETR1 of ETR1 is by of Arabidopsis by The The 1 The ETR1 by by The the 1 of the receptors of the subfamily 1 receptor ETR1 the subfamily receptor the the the of ETR1 The of the subfamily 1 receptors is the in in the ethylene receptors is the of of the ETR1 receptors that the the the of that the of receptor of The by the the the The ETR1 receptors of of The of the ethylene receptors the of the by by the The the a of a a of the ETR1 a of the is a that in a in the receptor that the receptor ethylene in a the receptor a that ethylene a of the a the that ETR1 of of receptors the of the ethylene receptor family ethylene receptors ethylene in a the of the the of by of of in the of is of the ethylene receptor family by ethylene in The of ETR1 ethylene ethylene ethylene of a ethylene that in a in the that a by the ethylene a of the receptor in ethylene the ethylene the by of receptor the receptor is ethylene ethylene receptors The the in a in that that of a the ethylene a that of the the the receptor a that of that the of is the of a receptor that the by that the ethylene receptor by ethylene in the in the The in receptor ethylene is the of the ethylene in of in the in the The the the ethylene The of in the in the of of in the the in the in the of The a by ethylene that ethylene receptors ethylene the in of the of ethylene the of of ethylene the receptors a in that the of ethylene The of the ethylene receptors is a that in a ethylene of ETR1 ethylene receptors that in that of ETR1 is of the of the of ETR1 the receptors is the of the in of ETR1 the of receptor of the receptors a that in of the of ethylene receptor in the of ETR1 the that a of the ethylene ETR1 by that the ethylene receptors is a that the ethylene receptors The receptors by of a in of ETR1 in in The of the in the of receptor the of receptor the that Arabidopsis ethylene receptors the in that in receptors that in that the of the receptors in a of the of the ethylene receptors of the receptors the of the ETR1 a the of is is that the the of of the ethylene receptors the in the ethylene receptors a Arabidopsis a in ETR1 ethylene a of The ethylene receptors ethylene ETR1 Arabidopsis ethylene is the The ethylene receptors the ethylene by receptor the ethylene by ethylene in The in a in of a in the receptor in a of receptors the of the the ethylene receptors the the receptors ethylene of that receptors subfamily 1 ethylene receptors of Arabidopsis a of subfamily receptors the receptors that the ethylene receptors the by in the receptors the The receptor a that ethylene is a the receptor ethylene the ETR1 ethylene by the that a receptor that ETR1 a ethylene The the of the receptor in the ethylene is in is ethylene by The receptor by receptors a of the receptor the of is the ethylene ethylene that of is the that is ethylene in of the receptor in subfamily 1 receptors The of receptors by is that that subfamily 1 receptors subfamily subfamily 1 ethylene receptors a ethylene of subfamily receptors the that subfamily 1 receptors ethylene in by subfamily a that the subfamily receptors a ethylene that subfamily 1 receptors ethylene by that subfamily 1 subfamily ethylene subfamily receptors subfamily 1 receptors subfamily receptors by ethylene that the subfamily 1 receptor ETR1 subfamily receptors that ethylene receptors of 1 in a that a the ethylene subfamily 1 subfamily is that subfamily 1 receptors a subfamily receptors The Arabidopsis The is a in a a a a in a by the the subfamily 1 receptors ETR1 the subfamily 1 receptors the the the of the subfamily receptors of of The hormone ethylene in in Arabidopsis ethylene of the of by ethylene of the of in five ethylene receptors in Arabidopsis The Arabidopsis receptors that ethylene in of the receptor the the The in The ethylene receptor family is ETR1 subfamily subfamily the The the ethylene receptors is a receptors is the of receptor a the is the that receptor a the in the the ethylene ethylene receptor that ethylene of a a of ethylene receptors of ethylene receptor a ethylene that ethylene receptors ethylene ethylene receptors ethylene is a of the ethylene receptors The of ethylene receptors in Arabidopsis by the of that ETR1 ethylene receptors in ETR1 the subfamily ethylene The ethylene receptors is a that the ethylene receptors in in a that the of The ethylene receptors the of ethylene in the of in the of the the a the the The the of the The the of the The the Arabidopsis The the Arabidopsis The the Arabidopsis The the of of a the the Arabidopsis The the of a the in the Arabidopsis by the The ethylene receptor receptor of the the the the the of the of Arabidopsis ethylene The of of Arabidopsis Arabidopsis Arabidopsis in of ethylene in in a 1 The the in by the of a of ETR1 the of the The a by the of a by the of the the a 1 a the of that the by the a a of the of of 1 1 The a The the by of ETR1 of in the The 1 The 1 The The The by of the ETR1 in of in the The the the of the the ETR1 ETR1 the the of of in the a The the of the The the The the of the the The the of of The the of the in by the ETR1 of the ETR1 the of ETR1 the of the in the The of the ETR1 by the the ETR1 a the ETR1 the of the the the of the a the of in by the The a the in a The the of the the a the the of that in the in of the the a the the The the of the The the of the The the Arabidopsis The the Arabidopsis The the Arabidopsis The the of of a the the Arabidopsis The the of a the in the Arabidopsis by the The ethylene receptor receptor of the the the the the of the of Arabidopsis ethylene The of of Arabidopsis Arabidopsis Arabidopsis in of ethylene in in a 1 The the in by the of a of ETR1 the of the The a by the of a by the of the the a 1 a the of that the by the a a of the of of 1 1 The a The the by of ETR1 of in the The 1 The 1 The The The by of the ETR1 in of in the The the the of the the ETR1 ETR1 the the of of in the a The the of the The the The the of the the The the of of The the of The the in by the ETR1 of the ETR1 the of ETR1 the of the in the The of the ETR1 by the the ETR1 a the ETR1 the of the the the of the a the of in by the The a the in a The the of the the a the the of that in the in ethylene receptors Arabidopsis the a the of the ethylene receptors The of ethylene receptor the of The ethylene receptor the the ethylene receptors The ethylene receptors the of the that the receptor the ethylene receptors by in the that ethylene receptors in ETR1 the ethylene receptors in The ethylene receptors by The the of the in of the ethylene ETR1 the ethylene that ETR1 that ETR1 the a of the that the of ETR1 by the ethylene the of the receptor the of ETR1 by that of the of ETR1 of the of ETR1 of the receptors in of the of the receptors The of ETR1 the ethylene receptors the of the subfamily ethylene receptors of the subfamily 1 member a of of ETR1 ethylene the ETR1 the ethylene a ethylene of is the by ethylene of in the of ethylene of ethylene by The of ETR1 the the of that of the of ETR1 ethylene the of ETR1 a the in The of ETR1 a the ethylene ethylene of the in a of ETR1 the in the of ethylene ETR1 of ETR1 ethylene that receptor in the of the of the receptor ethylene of ETR1 ethylene receptors in the of that in the of ethylene in the of the of ethylene of in the of ETR1 of The receptor ethylene is the in the is the ETR1 The the of ETR1 by the receptors is the the of by ETR1 in that ETR1 a that the that in of in the The in ethylene the of the five receptors that ethylene receptors in the of ethylene receptors ETR1 ethylene receptors a a receptor a of ETR1 a of the the of a the of the ETR1 the by of by the of a the the the of of of the of The of of the of of the in a a of the of a of the the the a subfamily 1 of the of ETR1 in the that the in that a the the of a is The of a ETR1 in the that ETR1 the ethylene receptors of of ETR1 in ETR1 by ETR1 ETR1 is by that ETR1 in the of ETR1 by the ETR1 a of by ETR1 by in ETR1 by the is by The the ETR1 the ETR1 the of the that the the of ETR1 by that the of ETR1 in Arabidopsis is by The that a of ETR1 the that in Arabidopsis ETR1 by the of the of ETR1 of ETR1 is by of Arabidopsis by The The 1 The ETR1 by by The the 1 of the receptors of the subfamily 1 receptor ETR1 the subfamily receptor the the the of ETR1 The of the subfamily 1 receptors is the in in the ethylene receptors is the of of the ETR1 receptors that the the the of that the of receptor of The by the the the The ETR1 receptors of of The of the ethylene receptors the of the by by the The the a of a a of the ETR1 a of the is a that in a in the receptor that the receptor ethylene in a the receptor a that ethylene a of the a the that ETR1 of of receptors the of the ethylene receptor family ethylene receptors ethylene in a the of the the of by of of in the of is of the ethylene receptor family by ethylene in The of ETR1 ethylene ethylene ethylene of a ethylene that in a in the that a by the ethylene a of the receptor in ethylene the ethylene the by of receptor the receptor is ethylene ethylene receptors The the in a in that that of a the ethylene a that of the the the receptor a that of that the of is the of a receptor that the by that the ethylene receptor by ethylene in the in the The in receptor ethylene is the of the ethylene in of in the in the The the the ethylene The of in the in the of of in the the in the in the of The a ethylene receptors Arabidopsis the a the of the ethylene receptors The of ethylene receptor the of The ethylene receptor the the ethylene receptors The ethylene receptors the of the that the receptor the ethylene receptors by in the ETR1 that ethylene receptors in ETR1 the ethylene receptors in The ethylene receptors by The the of the in of the ethylene ETR1 the ethylene The that ETR1 that ETR1 the a of the that the of ETR1 by the ethylene the of the receptor the of ETR1 by that of the of ETR1 of the of ETR1 of the receptors in of the of the receptors The of ETR1 the ethylene receptors the of the subfamily ethylene receptors of the subfamily 1 member a of of ETR1 ethylene the ETR1 the ethylene a ethylene of is the by ethylene of in the of ethylene of ethylene by The of ETR1 the the of that of the of ETR1 ethylene the of ETR1 a the in The of ETR1 a the ethylene ethylene of the in a of ETR1 the in the of ethylene ETR1 of ETR1 ethylene that receptor in the of the of the receptor ethylene of by ETR1 in that ETR1 a that the that in of in the The in ethylene the of the five receptors that ethylene receptors in the of ethylene receptors ETR1 ethylene receptors a a receptor a of ETR1 a of the the of a the of the ETR1 the by of by the of a the the the of of of the of The of of the of of the in a a of the of a of the the the a subfamily 1 of the the that the in that a the the of a is The of a ETR1 in the that ETR1 the ethylene receptors of of ETR1 in ETR1 by ETR1 ETR1 is by that ETR1 in the of ETR1 by the ETR1 a of by ETR1 by in ETR1 by the is by The the ETR1 the ETR1 the of the that the the of ETR1 by that the of ETR1 in Arabidopsis is by The that a of ETR1 the that in Arabidopsis ETR1 by the of the of ETR1 The of the receptors of the subfamily 1 receptor ETR1 the subfamily receptor the the the of ETR1 The of the subfamily 1 receptors is the in in the ethylene receptors is the of of the ETR1 receptors that the the the of that the of the is a that in a in the receptor that the receptor ethylene in a the receptor a that ethylene a of the a the that ETR1 of of receptors the of the ethylene receptor family ethylene receptors ethylene in a the of the the of by of of in the of is of the ethylene receptor family by ethylene in The of ETR1 ethylene ethylene ethylene of The a ethylene that in a in the that a by the ethylene a of the receptor in ethylene the ethylene the by of receptor the receptor is ethylene ethylene receptors The the in a in that that of a the ethylene a that of the the the receptor a that of that the of is the of a receptor that the by that the ethylene receptor by ethylene in the in the The in receptor ethylene is the by ethylene that ethylene receptors ethylene the in of the of ethylene the of of ethylene the receptors a in that the of ethylene The of the ethylene receptors is a that in a ethylene of ETR1 ethylene receptors that in that of ETR1 is of the of the of ETR1 the receptors is the of the in of ETR1 the of receptor of the receptors a that in of the of ethylene receptor in the of ETR1 the that a of the ethylene ETR1 by that the ethylene receptors is a that the ethylene receptors The receptors by of a in of ETR1 in in The of the in the of receptor the of receptor the that Arabidopsis ethylene receptors the in that in receptors that in that the of the receptors in a of the of the ethylene receptors of the receptors the of the ETR1 a the of is is that the the of of the ethylene receptors the in the ethylene receptors a Arabidopsis a in ETR1 ethylene a of The ethylene receptors ethylene ETR1 Arabidopsis ethylene is the The ethylene receptors the ethylene by receptor the ethylene by ethylene in The in a in of a in the receptor in a of receptors the of the the ethylene receptors the the receptors ethylene of that receptors subfamily 1 ethylene receptors of Arabidopsis a of subfamily receptors the receptors that the ethylene receptors the by in the receptors the The receptor a that ethylene is a the receptor ethylene the ETR1 ethylene by the that a receptor that ETR1 a ethylene The the of the receptor in the ethylene is in is ethylene by The receptor by receptors a of the receptor the of is the ethylene ethylene that of is the that is ethylene in of the receptor in subfamily 1 receptors The of receptors by is that that subfamily 1 receptors subfamily subfamily 1 ethylene receptors a ethylene of subfamily receptors the that subfamily 1 receptors ethylene in by subfamily a that the subfamily receptors a ethylene that subfamily 1 receptors ethylene by that subfamily 1 subfamily ethylene subfamily receptors subfamily 1 receptors subfamily receptors by ethylene that the subfamily 1 receptor ETR1 subfamily receptors that ethylene receptors of 1 in a that a the ethylene subfamily 1 subfamily is that subfamily 1 receptors a subfamily receptors The Arabidopsis The is a in a a a a in a by the the subfamily 1 receptors ETR1 the subfamily 1 receptors the the the of the subfamily receptors of of by ethylene that ethylene receptors ethylene the in of the of ethylene the of of ethylene the receptors a in that the of ethylene The of the ethylene receptors is a that in a ethylene of ETR1 ethylene receptors that in that of ETR1 is of the of the of ETR1 the receptors is the of the in of ETR1 the of receptor of the receptors a that in of the of ethylene receptor in the of ETR1 the that a of the ethylene The ETR1 by that the ethylene receptors is a that the ethylene receptors The receptors by of a in of ETR1 in in The of the in the of receptor the of receptor the that Arabidopsis ethylene receptors the in that in receptors that in that the of the receptors in a of the of the ethylene receptors of the receptors the of the ETR1 a the of is is that the the of of the ethylene receptors the in the ethylene receptors a Arabidopsis a in ETR1 ethylene a of The ethylene receptors ethylene ETR1 Arabidopsis ethylene is the The ethylene receptors the ethylene by receptor the ethylene by ethylene in The in a in of a in the receptor in a of receptors the of the the The ethylene receptors the the receptors ethylene of that receptors subfamily 1 ethylene receptors of Arabidopsis a of subfamily receptors the receptors that the ethylene receptors the by in the receptors the The receptor a that ethylene is a the receptor ethylene the ETR1 ethylene by the that a receptor that ETR1 a ethylene The the of the receptor in the ethylene is in is ethylene by The receptor by receptors a of the receptor the of is the ethylene ethylene that of is the that is ethylene in of the receptor in subfamily 1 receptors The of receptors by is that that subfamily 1 receptors subfamily subfamily 1 ethylene receptors a ethylene of subfamily receptors the that subfamily 1 receptors ethylene in by subfamily a that the subfamily receptors a ethylene that subfamily 1 receptors ethylene by that subfamily 1 subfamily ethylene subfamily receptors subfamily 1 receptors subfamily receptors by ethylene The that the subfamily 1 receptor ETR1 subfamily receptors that ethylene receptors of 1 in a that a the ethylene subfamily 1 subfamily is that subfamily 1 receptors a subfamily receptors The Arabidopsis The is a in a a a a in a by the the subfamily 1 receptors ETR1 the subfamily 1 receptors the the the of the subfamily receptors of of the the the the the