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Satoshi Ohdachi, Toshiki Aoi, Food Habits of Brown Bears in Hokkaido, Japan, Bears: Their Biology and Management, Vol. 7, A Selection of Papers from the Seventh International Conference on Bear Research and Management, Williamsburg, Virginia, USA, and Plitvice Lakes, Yugoslavia, February and March 1986 (1987), pp. 215-220
Ohdachi et al. (Thu,) studied this question.