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Journal Article Landscape Ecology Get access Dean L. Urban, Dean L. Urban Search for other works by this author on: Oxford Academic Google Scholar Robert V. O'Neill, Robert V. O'Neill Search for other works by this author on: Oxford Academic Google Scholar Herman H. Shugart, Jr. Herman H. Shugart, Jr. Search for other works by this author on: Oxford Academic Google Scholar BioScience, Volume 37, Issue 2, February 1987, Pages 119–127, https://doi.org/10.2307/1310366 Published: 01 February 1987
Urban et al. (Sun,) studied this question.