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Since 1951, the biomass of macrozooplankton in waters off southern California has decreased by 80 percent. During the same period, the surface layer warmed-by more than 1.5 degrees C in some places-and the temperature difference across the thermocline increased. Increased stratification resulted in less lifting of the thermocline by wind-driven upwelling. A shallower source of upwelled waters provided less inorganic nutrient for new biological production and hence supported a smaller zooplankton population. Continued warming could lead to further decline of zooplankton.
Roemmich et al. (Fri,) studied this question.