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R. Scott Russell, D. Ward Cutler, S. E. Jacobs, Alexander King, A. G. Pollard, Physiological and Ecological Studies on an Arctic Vegetation: II. The Development of Vegetation in Relation to Nitrogen Supply and Soil Micro- Organisms on Jan Mayen Island, Journal of Ecology, Vol. 28, No. 2 (Aug., 1940), pp. 269-288
Russell et al. (Thu,) studied this question.