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John Okuley, Jonathan Lightner, Kenneth Feldmann, Narendra Yadav, Ellen Lark, John Browse, Arabidopsis FAD2 Gene Encodes the Enzyme That Is Essential for Polyunsaturated Lipid Synthesis, The Plant Cell, Vol. 6, No. 1 (Jan., 1994), pp. 147-158
Okuley et al. (Sat,) studied this question.