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W. Ronald Heyer, A. Stanley Rand, Carlos Alberto Goncalvez da Cruz, Oswaldo L. Peixoto, Decimations, Extinctions, and Colonizations of Frog Populations in Southeast Brazil and Their Evolutionary Implications, Biotropica, Vol. 20, No. 3 (Sep., 1988), pp. 230-235
Heyer et al. (Thu,) studied this question.
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