Spatial heterogeneity in urban ecosystems: reconceptualizing land cover and a framework for classification

Urban areas are heterogeneous. Transitions in architecture and building density, vegetation, economic activity, and culture can occur at the scale of city blocks. Ecologists have been criticized for treating the city as homogeneous and urbanization as one-dimensional. To develop ecological understanding of integrated human–natural systems, the fine-scale heterogeneity of their built and natural components must be quantified. There have been calls for the integration of the biophysical and human components of systems, but here we provide a new tool to quantify this integrated heterogeneity by reconceptualizing urban land-use and land-cover classification approaches. This new tool, High Ecological Resolution Classification for Urban Landscapes and Environmental Systems (HERCULES), balances detail and efficiency and is flexible, allowing it to be used for interdisciplinary research, with ancillary datasets, and across urban systems.