Key points are not available for this paper at this time.
Harry Hoogstraal, Frederick H. S. Roberts, Glen M. Kohls, Vernon J. Tipton, Review of Haemaphysalis (Kaiseriana) longicornis Neumann (Resurrected) of Australia, New Zealand, New Caledonia, Fiji, Japan, Korea, and Northeastern China and USSR, and Its Parthenogenetic and Bisexual Populations (Ixodoidea, Ixodidae), The Journal of Parasitology, Vol. 54, No. 6 (Dec., 1968), pp. 1197-1213
Hoogstraal et al. (Sun,) studied this question.