Spatial simulation of forest succession and timber harvesting using LANDIS

The LANDIS model simulates ecological dynamics, including forest succession, disturbance, seed dispersal and establishment, fire and wind disturbance, and their interactions. We describe the addition to LANDIS of capabilities to simulate forest vegetation management, including harvest. Stands (groups of cells) are prioritized for harvest using one of four ranking algorithms that use criteria related to forest management objectives. Cells within a selected stand are harvested according to the species and age cohort removal rules specified in a prescription. These flexible removal rules allow simulation of a wide range of prescriptions such as prescribed burning, thinning, single-tree selection, and clear-cutting. We present a case study of the application of LANDIS to a managed watershed in the Missouri (U.S.A.) Ozark Mountains to illustrate the utility of this approach to simulate succession as a response to forest management and other disturbance. The different cutting practices produced differences in species and size-class composition, average patch sizes (for patches defined by forest type or by size class), and amount of forest edge across the landscape. The capabilities of LANDIS provide a modeling tool to investigate questions of how timber management changes forest composition and spatial pattern, providing insight into ecological response to forest management.