Key points are not available for this paper at this time.
H. V. Cornell, J. H. Lawton, Species Interactions, Local and Regional Processes, and Limits to the Richness of Ecological Communities: A Theoretical Perspective, Journal of Animal Ecology, Vol. 61, No. 1 (Feb., 1992), pp. 1-12
Cornell et al. (Sat,) studied this question.