What question did this study set out to answer?

To review the East Asian species of Rheocricotopus and describe new species in the subgenus Psilocricotopus.

February 2, 2026European Journal of Taxonomy0 citationsOpen Access

Taxonomic review of Rheocricotopus Brundin, 1956 (Diptera: Chironomidae: Orthocladiinae) from East Asia, with descriptions of twelve new species

HNHiromi NiitsumaIndustrial Research Institute of Shizuoka Prefecture HTHongqu TangJinan University

Key Points

To review the East Asian species of Rheocricotopus and describe new species in the subgenus Psilocricotopus.
Illustration and description of twelve new species.
Identification of additional life stages for previously known species.
Application of taxonomic keys for classification.
Twelve new species were described and illustrated.
Total of 28 new synonyms were established.
Revisions to generic and subgeneric diagnoses to accommodate new findings.

Abstract

East Asian species of Rheocricotopus are reviewed and 12 species are illustrated and described as new to science in the subgenus Psilocricotopus Sæther: R. (P.) albulus sp. nov., R. (P.) bannaensis sp. nov., R. (P.) beibengensis sp. nov., R. (P.) binotatus sp. nov., R. (P.) chinogodavarius sp. nov., R. (P.) fasciolatus sp. nov., R. (P.) hanmiensis sp. nov., R. (P.) masarui sp. nov., R. (P.) monovittatus sp. nov., R. (P.) nippobifasciatus sp. nov., R. (P.) protuberans sp. nov., and R. (P.) yunnanensis sp. nov. Additionally, the female adult, pupa and larva of R. (P.) togakuroasi (Sasa R. chalybeatus bicoloratus Caspers R. akagisecundus Kikuchi R. baishanensis Wang R. reduncusoides Namayandeh R. kamimonji Sasa Parorthocladius negoroi Yamamoto, 2011 and Orthocladius piloculatus Kobayashi, 2012 are treated as junior synonyms of Parorthocladius intermedius (Tokunaga, 1939) comb. nov.; R. tobatervicesimus Kikuchi & Sasa, 1990, Paratrichocladius unabrevis Sasa, 1996 and R. rotundus Liu, Lin & Wang, 2014 are treated as junior synonyms of Paracricotopus irregularis Niitsuma, 1990. In addition, two new combinations, Parorthocladius intermedius (Tokunaga, 1939) comb. nov. and Paracricotopus brachypus (Wang & Zheng, 1991) comb. nov. are proposed here. To accommodate these species, generic and subgeneric diagnoses are emended. A key to the known male adults from East Asia is provided.

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