Key points are not available for this paper at this time.
R. E. Comstock, H. F. Robinson, The Components of Genetic Variance in Populations of Biparental Progenies and Their Use in Estimating the Average Degree of Dominance, Biometrics, Vol. 4, No. 4 (Dec., 1948), pp. 254-266
Comstock et al. (Wed,) studied this question.