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We describe a general framework for understanding the ecological processes that operate at landscape scales. The composition of habitat types in a landscape and the physiognomic or spatial arrangement of those habitats are the two essential features that are required to describe any landscape. As such, these two features affect four basic ecological processes that can influence population dynamics or community structure. The first two of these processes, landscape complementation and landscape supplementation, occur when individuals move between patches in the landscape to make use of nonsubstitutable and substitutable resources, respectively
Dunning et al. (Thu,) studied this question.
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