Observational analysis identified key aspects of breeding behaviour and communication during nesting, highlighting the significance of display patterns.
The breeding cycle begins in late September, when the mature males come ashore after their winter at sea to occupy their nesting sites, and ends in early April, when the breeders return to sea following their annual moult. This cycle of nesting and moult is outlined and the main patterns in a complex sign-language of display and posturing are described.
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John Warham (1971) studied this question.
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