Observational analysis shows different crown fade rates in douglas-fir trees due to varying weather patterns, indicating significant environmental influence.
Foliage of Douglas-fir trees, observed for two seasons following lethal attack by the Douglas-fir beetle turned color at different rates depending upon weather patterns. In 1964, a cool and wet year, some killed trees faded in July, but most did not turn until after May 1965 and continued until that fall. In 1965, a warmer and drier year, 70 percent of the trees attacked in the spring were red by October as compared to 25 percent in the same month a year previously. By July 1966, all trees attacked the previous year were bare.
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Belluschi et al. (1969) studied this question.
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