The first fossil of a tenebrionoid taxonomic enigma: Agnathus Germar (Coleoptera: Pyrochroidae: Agnathinae) in Bitterfeld amber, with remarks about age and geographic origin of the fossil

Based on an inclusion in Bitterfeld amber, the first extinct species of the subfamily Agnathinae (Pyrochroidae) is described and illustrated. Agnathus groehni sp. nov. differs from two extant congeners in having the smaller body size; transverse W-shaped groove near posterior pronotal margin; the unicolourous black body and the elytral pubescence uniform without tomentose pattern. As demonstrated herein, the genus Agnathus is morphologically stable over at least 23 Ma years and inhabited the Western Palaearctic already in the Palaeogene. Zoogeography of extant species, palaeobionomy of the extinct species and difficulties in its reconstruction, as well as age and origin of Bitterfeld amber, are briefly discussed.