State-dependent stiffness enhances wave propagation along elastic filaments

We study an elastic filament beating in a viscous fluid with curvature-dependent bending stiffness. Our numerical and experimental investigations reveal that such differential stiffness can sustain planar bending waves far along flexible filaments. In particular, basal actuation is a viable, parsimonious mechanism for generating high-amplitude planar bending waves, in stark contrast to the uniform-stiffness case. Further, the resulting beat patterns closely resemble the power-and-recovery strokes of propulsive biological filaments, suggesting applications in robotic and engineered systems.