W. Volkl, H. Zwolfer, M. Romstock-Volkl, C. Schmelzer, Habitat Management in Calcareous Grasslands: Effects on the Insect Community Developing in Flower Heads of Cynarea, Journal of Applied Ecology, Vol. 30, No. 2 (1993), pp. 307-315
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Völkl et al. (1993) studied this question.
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