Mark A. Keech, R. Terry Bowyer, Jay M. Ver Hoef, Rodney D. Boertje, Bruce W. Dale, Thomas R. Stephenson, Life-History Consequences of Maternal Condition in Alaskan Moose, The Journal of Wildlife Management, Vol. 64, No. 2 (Apr., 2000), pp. 450-462
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Keech et al. (2000) studied this question.
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