Daniel H. Pletscher, Robert R. Ream, Diane K. Boyd, Michael W. Fairchild, Kyran E. Kunkel, Population Dynamics of a Recolonizing Wolf Population, The Journal of Wildlife Management, Vol. 61, No. 2 (Apr., 1997), pp. 459-465
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Pletscher et al. (1997) studied this question.
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