Key points are not available for this paper at this time.
Daniel H. Pletscher, Robert R. Ream, Diane K. Boyd, Michael W. Fairchild, Kyran E. Kunkel, Population Dynamics of a Recolonizing Wolf Population, The Journal of Wildlife Management, Vol. 61, No. 2 (Apr., 1997), pp. 459-465
Pletscher et al. (Tue,) studied this question.