R. Edward DeWalt, Kenneth W. Stewart, Stephen R. Moulton II, James H. Kennedy, Summer Emergence of Mayflies, Stoneflies, and Caddisflies from a Colorado Mountain Stream, The Southwestern Naturalist, Vol. 39, No. 3 (Sep., 1994), pp. 249-256
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DeWalt et al. (1994) studied this question.
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