Cynthia P. de Almeida Prado, Masao Uetanabaro, Frederico S. Lopes, Reproductive Strategies of Leptodactylus chaquensis and L. podicipinus in the Pantanal, Brazil, Journal of Herpetology, Vol. 34, No. 1 (Mar., 2000), pp. 135-139
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Prado et al. (2000) studied this question.