The (13)C (12)C isotope ratios and the leaf anatomy of 18 species and varieties of Euphorbia native to the Hawaian Islands indicated that all possess C(4) photosynthesis. These species range from small prostrate coastal strand shrubs to shrubs and trees in rainforest and bog habitats. The results show that C(4) photosynthesis occurs in plants from a much wider range of habitats and life-forms than has been previously reported.
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Pearcy et al. (1975) studied this question.
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