D. Megan Helfgott, Javier Francisco-Ortega, Arnoldo Santos-Guerra, Robert K. Jansen, Beryl B. Simpson, Biogeography and Breeding System Evolution of the Woody Bencomia Alliance (Rosaceae) in Macaronesia Based on ITS Sequence Data, Systematic Botany, Vol. 25, No. 1 (Jan. - Mar., 2000), pp. 82-97
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Helfgott et al. (2000) studied this question.
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