W. Nelson Beyer, Donald M. Saari, Activity and Ecological Distribution of the Slug, Arion subfuscus (Draparnaud) (Stylommatophora, Arionidae), The American Midland Naturalist, Vol. 100, No. 2 (Oct., 1978), pp. 359-367
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Beyer et al. (1978) studied this question.
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