Xinnian Dong, Michael Mindrinos, Keith R. Davis, Frederick M. Ausubel, Induction of Arabidopsis Defense Genes by Virulent and Avirulent Pseudomonas syringae Strains and by a Cloned Avirulence Gene, The Plant Cell, Vol. 3, No. 1 (Jan., 1991), pp. 61-72
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Dong et al. (1991) studied this question.