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Wilhelm Fortelius, Rainer Rosengren, Daniel Cherix, Dominique Chautems, Queen Recruitment in a Highly Polygynous Supercolony of Formica lugubris (Hymenoptera, Formicidae), Oikos, Vol. 67, No. 2 (Jun., 1993), pp. 193-200
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Fortelius et al. (1993) studied this question.
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