Quantitative analyses of crude protein, crude fat, crude fiber, ash, calcium, phosphorus, and carotene were performed on 9 forage species, collected on 5 dates during a year. Crude protein levels were low in most forage species by mid-winter. Phosphorus levels were low in most plants throughout the year. Lichens, an important winter forage of reindeer, appear to be deficient in crude protein, calcium, phosphorus, and carotene.
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George W. Scotter (1972) studied this question.