Mauro Galetti, Alexine Keuroghlian, Lais Hanada, Maria Inez Morato, Frugivory and Seed Dispersal by the Lowland Tapir (Tapirus terrestris) in Southeast Brazil, Biotropica, Vol. 33, No. 4 (Dec., 2001), pp. 723-726
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Galetti et al. (2001) studied this question.
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