A. C. Stevenson, P. D. Moore, Studies in the Vegetational History of S. W. Spain. IV. Palynological Investigations of a Valley Mire at El Acebron, Huelva, Journal of Biogeography, Vol. 15, No. 2 (Mar., 1988), pp. 339-361
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Stevenson et al. (1988) studied this question.