The forest stand structure class “old-growth” has previously been qualitatively described as having several distinct “sub-structures.” Species composition, diameter distribution, and other structural features commonly associated with old-growth in the Pacific Northwest are quite variable. We determined which quantitative stand structure variables are commonly found together using the Spearman correlation and non-metric multidimensional analysis. Some features were more commonly found together than others, indicating different old-growth stand types, or sub-structures. Cluster analysis classified the old-growth forests into four groups: Douglas-fir dominance, shade tolerant species dominance, and intermediate groups. The intermediate groups were split by the density of large logs and large shade tolerant trees. The old-growth sub-structures appear to change from one to another as the old forest develops.
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Park et al. (2015) studied this question.
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