Four pigeons were trained to peck at either of two response-keys. Pecking at either key occasionally produced a secondary reinforcer. Then, in the presence of the secondary reinforcer, further pecking occasionally produced the primary reinforcer, food. The relative rate at which each pigeon pecked to obtain a secondary reinforcer equalled the relative rate of primary reinforcement in its presence.
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R. J. Herrnstein (1964) studied this question.
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