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Katherine J. Elliott, James M. Vose, Wayne T. Swank, Paul V. Bolstad, Long-Term Patterns in Vegetation-Site Relationships in a Southern Appalachian Forest, The Journal of the Torrey Botanical Society, Vol. 126, No. 4 (Oct. - Dec., 1999), pp. 320-334
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Elliott et al. (1999) studied this question.
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