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Chicita F. Culberson, William Louis Culberson, Debra A. Arwood, Physiography and Fumarprotocetraric Acid Production in the Cladonia chlorophaea Group in North Carolina, The Bryologist, Vol. 80, No. 1 (Spring, 1977), pp. 71-75
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Culberson et al. (1977) studied this question.
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