J. C. Z. Woinarski, P. J. Whitehead, D. M. J. S. Bowman, J. Russell-Smith, Conservation of Mobile Species in a Variable Environment: The Problem of Reserve Design in the Northern Territory, Australia, Global Ecology and Biogeography Letters, Vol. 2, No. 1 (Jan., 1992), pp. 1-10
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Woinarski et al. (1992) studied this question.
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