Key points are not available for this paper at this time.
Jacek M. Szymura, Nicholas H. Barton, Genetic Analysis of a Hybrid Zone Between the Fire-Bellied Toads, Bombina bombina and B. variegata, Near Cracow in Southern Poland, Evolution, Vol. 40, No. 6 (Nov., 1986), pp. 1141-1159
Szymura et al. (Sat,) studied this question.