B. A. Hutchison, D. R. Matt, R. T. McMillen, L. J. Gross, S. J. Tajchman, J. M. Norman, The Architecture of a Deciduous Forest Canopy in Eastern Tennessee, U.S.A., Journal of Ecology, Vol. 74, No. 3 (Sep., 1986), pp. 635-646
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Hutchison et al. (1986) studied this question.
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