N. P. R. Anteb, R. Hernandez, E. M. Medina, The Photosynthetic Capacity and Leaf Nitrogen Concentration as Related to Light Regime in Shade Leaves of a Montane Tropical Forest Tree, Tetrochidium rubrivenium, Functional Ecology, Vol. 10, No. 4 (Aug., 1996), pp. 491-500
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Anteb et al. (1996) studied this question.
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