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Ola Inghe, Carl Olof Tamm, Survival and Flowering of Perennial Herbs. IV. The Behaviour of Hepatica Nobilis and Sanicula Europaea on Permanent Plots during 1943-1981, Oikos, Vol. 45, No. 3 (Dec., 1985), pp. 400-420
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Inghe et al. (1985) studied this question.
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