Mogens G. Nielsen, Torben F. Jensen, Ib Holm-Jensen, Effect of Load Carriage on the Respiratory Metabolism of Running Worker Ants of Camponotus herculeanus (Formicidae), Oikos, Vol. 39, No. 2 (Aug., 1982), pp. 137-142
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Nielsen et al. (1982) studied this question.
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