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A.W. Illius, S.D. Albon, J.M. Pemberton, I.J. Gordon, T.H. Clutton-Brock, Selection for Foraging Efficiency During a Population Crash in Soay Sheep, Journal of Animal Ecology, Vol. 64, No. 4 (Jul., 1995), pp. 481-492
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Illius et al. (1995) studied this question.
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