David R. Breininger, Paul A. Schmalzer, C. Ross Hinkle, Estimating Occupancy of Gopher Tortoise (Gopherus polyphemus) Burrows in Coastal Scrub and Slash Pine Flatwoods, Journal of Herpetology, Vol. 25, No. 3 (Sep., 1991), pp. 317-321
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Breininger et al. (1991) studied this question.
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