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Edward J. Heist, John E. Graves, John A. Musick, Population Genetics of the Sandbar Shark (Carcharhinus plumbeus) in the Gulf of Mexico and Mid-Atlantic Bight, Copeia, Vol. 1995, No. 3 (Aug. 18, 1995), pp. 555-562
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Heist et al. (1995) studied this question.
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