W. F. Morris, M. V. Price, N. M. Waser, J. D. Thomson, B. Thomson, D. A. Stratton, Systematic Increase in Pollen Carryover and Its Consequences for Geitonogamy in Plant Populations, Oikos, Vol. 71, No. 3 (Dec., 1994), pp. 431-440
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Morris et al. (1994) studied this question.
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