N. I. Klomp, R. W. Furness, Non-Breeders as a Buffer Against Environmental Stress: Declines in Numbers of Great Skuas on Foula, Shetland, and Prediction of Future Recruitment, Journal of Applied Ecology, Vol. 29, No. 2 (1992), pp. 341-348
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Klomp et al. (1992) studied this question.
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