Paul T. Chippindale, Andrew H. Price, John J. Wiens, David M. Hillis, Phylogenetic Relationships and Systematic Revision of Central Texas Hemidactyliine Plethodontid Salamanders, Herpetological Monographs, Vol. 14 (2000), pp. 1-80
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Chippindale et al. (2000) studied this question.
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