E. M. Goolish, R. S. Burton, Energetics of Osmoregulation in an Intertidal Copepod: Effects of Anoxia and lipid Reserves on the Pattern of Free Amino Accumulation, Functional Ecology, Vol. 3, No. 1 (1989), pp. 81-89
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Goolish et al. (1989) studied this question.
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