John D. McEachran, Katherine A. Dunn, Phylogenetic Analysis of Skates, a Morphologically Conservative Clade of Elasmobranchs (Chondrichthyes: Rajidae), Copeia, Vol. 1998, No. 2 (May 1, 1998), pp. 271-290
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McEachran et al. (1998) studied this question.
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