David S. Barrington, Christopher H. Haufler, Charles R. Werth, Hybridization, Reticulation, and Species Concepts in the Ferns, American Fern Journal, Vol. 79, No. 2, Species Concepts in Pteridophytes (Apr. - Jun., 1989), pp. 55-64
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Barrington et al. (1989) studied this question.
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